Satyrini

Satyrini
	Oranje zandoogje (Pyronia tithonus)
Taxonomische indeling
Rijk:	Animalia (Dieren)
Stam:	Arthropoda (Geleedpotigen)
Klasse:	Insecta (Insecten)
Orde:	Lepidoptera (Vlinders)
Superfamilie:	Papilionoidea
Familie:	Nymphalidae
Onderfamilie:	Satyrinae
Geslachtengroep
	Satyrini; Boisduval, 1833
	Afbeeldingen op Wikimedia Commons
	Satyrini op Wikispecies
Portaal	Biologie ; Insecten

De Satyrinii zijn een geslachtengroep van vlinders in de onderfamilie Satyrinae van de familie Nymphalidae. De hier gegeven onderverdeling in subtribus en geslachten volgt die van Peña et al. van 2006,^[1] aangevuld uit het overzicht van geslachten door de Nymphalidae Systematics Group.^[2]

Subtribus en geslachten

Parargina Tutt, 1896

Pararge Hübner, 1819
Chonala Moore, 1893*^[3]
Kirinia Moore, 1893
- = Esperarge Nekrutenko, 1988
- = Esperia Nekrutenko, 1987 non Esperia Hübner, 1825 (Oecophoridae)
- = Esperella Nekrutenko, 1987 non Esperella Vosmaer, 1887 (Demospongiae)
Lasiommata Westwood, 1841
- = Amecera Butler, 1867
Lopinga Moore, 1893
- = Crebeta Moore, 1893
- = Polyargia Verity, 1957
Nosea Koiwaya, 1993*
Orinoma Gray, 1846*
Rhaphicera Butler, 1867*
Tatinga Moore, 1893*

Lethina Reuter, 1896

Lethe Hübner, 1819
- = Tanaoptera Billberg, 1820
- = Zophoessa Doubleday, 1849
- = Debis Doubleday, 1849
- = Charma Doherty, 1886
- = Tansima Moore, 1881
- = Rangbia Moore, 1892
- = Nemetis Moore, 1892
- = Pegada Moore, 1892
- = Kirrodesa Moore, 1892
- = Placilla Moore, 1892
- = Archondesa Moore, 1892
- = Choranesa Moore, 1892
- = Dionana Moore, 1892
- = Sinchula Moore, 1892
- = Kerrata Moore, 1892
- = Putlia Moore, 1892
- = Harima Moore, 1892
- = Magula Fruhstorfer, 1911
- = Hermias Fruhstorfer, 1911
Aphysoneura Karsch, 1894*
- = Rhaphiceropsis Sharpe, 1894
Enodia Hübner, 1819
Hanipha Moore, 1880*
Mandarinia Leech, 1892*
Neope Moore, 1866
- = Enope Moore, 1857 non Enope Walker, 1854 (Erebidae)
- = Neope Butler, 1867
- = Blanaida Kirby, 1871
- = Patala Moore, 1892
† Neorinella Martins Neto et al., 1993*
† Neorinopsis Butler, 1873*
Ninguta Moore, 1892*
- = Aranda Fruhstorfer, 1909
† Pseudoneorina Nel & Descimon, 1986*
Ptychandra C. & R. Felder, 1861*
† Satyrites Scudder, 1872*
- = Lethites Scudder, 1875
Satyrodes Scudder, 1875
- = Argus Scopoli, 1777

Mycalesina Reuter, 1896

Mycalesis Hübner, 1818
Bicyclus Kirby, 1871
Bletogona C. & R. Felder, 1867*
Brakefieldia Aduse-Poku, Lees & Wahlberg, 2016
Hallelesis Condamin, 1961
Heteropsis Westwood, 1850*
Nirvanopsis Vane-Wright, 2003*
Orsotriaena Wallengren, 1858^[4]
Pseudomycalesis Tsukada & Nishiyama, 1979*
- voor synoniemen, zie de subtribus

Eritina Miller, 1968

Erites Westwood, 1851*
Coelites Westwood, 1850*

Ragadiina Herrich-Schaeffer, 1864

Ragadia Westwood, 1851*
Acrophtalmia C. & R. Felder, 1861*
Acropolis Hemming, 1934*
- = Pharia Fruhstorfer, 1911 non Pharia Gray, 1840 (Asteroidea)

Coenonymphina Tutt, 1896

Coenonympha Hübner, 1819
- = Chortobius Dunning & Pickard, 1858
- = Sicca Verity, 1953
Lyela Swinhoe, 1908*
- = Dubierebia Muschamp, 1915
Sinonympha Lee, 1974*
Triphysa Zeller, 1850*
- = Phryne Herrich-Schäffer, 1844

Hypocystina Miller, 1968

Hypocysta Westwood, 1851
Altiapa Parsons, 1986*
Argynnina Butler, 1867*
Argyronympha Mathew, 1886
Argyrophenga Doubleday, 1845
Dodonidia Butler, 1884
Erebiola Fereday, 1879
Erycinidia Rothschild & Jordan, 1905*
- = Pieridopsis Rothschild & Jordan, 1905
Geitoneura Butler, 1867
Harsiesis Fruhstorfer, 1911*
Heteronympha Wallengren, 1858
- = Hipparchioides Butler, 1867
Hyalodia Jordan, 1924*
Lamprolenis Godman & Salvin, 1881
Nesoxenica Waterhouse & Lyell, 1914
- = Xeniconympha Novicki, 1923
Oreixenica Waterhouse & Lyell, 1914
Paratisiphone Watkins, 1928*
Percnodaimon Butler, 1876
Platypthima Rothschild & Jordan, 1905*
Tisiphone Hübner, 1819
- = Xenica Westwood, 1851
Zipaetis Hewitson, 1863

Euptychiina Miller, 1968

Euptychia Hübner, 1818
Amphidecta Butler, 1867
Archeuptychia Forster, 1964*
Caenoptychia Le Cerf, 1919*
- = Ristia Gagarin, 1936
Caeruleuptychia Forster, 1964
- = Weymerana Forster, 1964
Capronnieria Forster, 1964*
Cepheuptychia Forster, 1964
Cercyeuptychia Miller & Emmel, 1971*
Chloreuptychia Forster, 1964
Cissia Doubleday, 1848
- = Argyreuptychia Forster, 1964
- = Vareuptychia Forster, 1964
Coeruleotaygetis Forster, 1964*
Cyllopsis Felder, 1869
Erichthodes Forster, 1964
Euptychoides Forster, 1964
Forsterinaria Gray, 1973
- = Haywardina Forster, 1964 non Haywardina Aczel, 1952 (Diptera)
- = Forsterina Lamas, 1977 non Forsterina Lehtinen, 1967 (Araneae)
Godartiana Forster, 1964
Guainanaza Freitas & Peña, 2006*
Harjesia Forster, 1964
Hermeuptychia Forster, 1964
Magneuptychia Forster, 1964
Megeuptychia Forster, 1964*
Megisto Hübner, 1819*
Moneuptychia Forster, 1964
- = Carminda Ebert & Dias, 1997
Neonympha Hübner, 1818
Oressinoma Doubleday, 1849^[4]
- = Ocalis Westwood, 1851
- = Ocalis Boisduval, 1870
Palaeonympha Butler, 1871
Paramacera Butler, 1868
Parataygetis Forster, 1964
Pareuptychia Forster, 1964
Paryphthimoides Forster, 1964
Pharneuptychia Forster, 1964
Pindis Felder, 1869
Posttaygetis Forster, 1964
Praefaunula Forster, 1964*
Pseudeuptychia Forster, 1964*
Pseudodebis Forster, 1964*
Rareuptychia Forster, 1964
Satyrotaygetis Forster, 1964*
Splendeuptychia Forster, 1964
Taydebis Freitas, 2003*
Taygetina Forster, 1964*
Taygetis Hübner, 1819
Taygetomorpha Miller, 2004*
Yphthimoides Forster, 1964
Zischkaia Forster, 1964

Erebiina Tutt, 1896

Erebia Dalman, 1816
- = Syngea Hübner, 1819
- = Epigea Hübner, 1819
- = Phorcis Hübner, 1819
- = Marica Hübner, 1819
- = Gorgo Hübner, 1819
- = Triariia Verity, 1953
- = Truncaefalcia Verity, 1953
- = Medusia Verity, 1953
- = Simplicia Verity, 1953
- = Atercoloratus Bang-Haas, 1938

Maniolina Grote, 1897

Maniola Schrank, 1801
- = Epinephele Hübner, 1819
Aphantopus Wallengren, 1853
Cercyonis Scudder, 1875
Hyponephele Muschamp, 1915
Proterebia Roos & Arnschied, 1980
Pyronia Hübner, 1819
- = Idata de Lesse, 1952
- = Pasiphana de Lesse, 1952

Pronophilina Reuter, 1896

Pronophila Doubleday, 1849
- = Mygona Westwood, 1851
Altopedaliodes Forster, 1964
Antopedaliodes Forster, 1964
Apexacuta Pyrcz, 2004
Argyrophorus Blanchard, 1852
Arhuaco Adams & Bernard, 1977
Auca Hayward, 1953
Calisto Hübner, 1823
Cheimas Thieme, 1907
Chillanella Herrera, 1966
Corades Hewitson, 1849
- = Corades Doubleday, 1848 (nomen nudum)
- = Panarche Thieme, 1907
Corderopedaliodes Forster, 1964
Cosmosatyrus C. & R. Felder, 1867
Daedalma Hewitson, 1858
Dangond Adams & Bernard, 1979
Diaphanos Adams & Bernard, 1981
Drucina Butler, 1872
Druphila Pyrcz, 2004
Elina Blanchard, 1852
Eretris Thieme, 1905
Etcheverrius Herrera, 1965
Eteona Doubleday, 1848
Faunula C. & R. Felder, 1867
Foetterleia Viloria, 2004
- = Foetterleia Viloria, 2003 (nom. nud.)
Gyrocheilus Butler, 1867
Haywardella Herrera, 1966
Homoeonympha C. & R. Felder, 1867
- = Stygnus Felder & Felder, 1867 non Stygnus Perty, 1833 (Opiliones)
- = Stygnolepis Strand, 1942
- = Erebina Bryk, 1944
Ianussiusa Pyrcz & Viloria, 2004
Idioneurula Strand, 1932
- = Idioneura Felder & Felder, 1867 non Idioneura Selys, 1860 (Odonata)
Junea Hemming, 1964
- = Polymastus Thieme, 1907
Lasiophila C. & R. Felder, 1859
Lymanopoda Westwood, 1851
- = Sarromia Westwood, 1851
- = Zabirnia Hewitson, 1877
- = Trophonina Röber, 1889
Manerebia Staudinger, 1897
- = Penrosada Brown, 1944
- = Posteuptychia Forster, 1964
Mygona Thieme, 1907
Nelia Hayward, 1953
Neomaenas Wallengren, 1858
- = Stibomorpha Butler, 1874
Neomaniola Hayward, 1949
- = Pseudomaniola Weymer, 1890
Neopedaliodes Viloria, Miller & Miller, 2004
- = Neopedaliodes Viloria, 2002 (nom. nud.)
Neosatyrus Wallengren, 1858
Oxeoschistus Butler, 1867
- = Dioriste Thieme, 1907
Palmaris Herrera, 1965
Pampasatyrus Hayward, 1953
- = Pseudocercyonis Miller & Emmel, 1971
Pamperis Heimlich, 1959
Panyapedaliodes Forster, 1964
- = Muscopedaliodes Forster, 1964
Paramo Adams & Bernard, 1977
Parapedaliodes Forster, 1964
Pedaliodes Butler, 1867
Pherepedaliodes Forster, 1964
Physcopedaliodes Forster, 1964
Praepedaliodes Forster, 1964
Praepronophila Forster, 1964
Proboscis Thieme, 1907
Protopedaliodes Viloria & Pyrcz, 1994
Pseudomaniola Weymer, 1890
- = Catargynnis Röber, 1892
Punapedaliodes Forster, 1964
Punargentus Heimlich, 1963
Quilaphoetosus Herrera, 1966
Redonda Adams & Bernard, 1981
Sabatoga Staudinger, 1897
- = Zapatoca Apolinar, 1924
Sierrasteroma Adams & Bernard, 1977
Spinantenna Hayward, 1953
Steremnia Thieme, 1905
- = Pseudosteroma Weymer, 1912
Steroma Westwood, 1850
Steromapedaliodes Forster, 1964
- = Steropoda Thieme, 1905
Stuardosatyrus Herrera & Etcheverry, 1965
Tamania Prycz, 1995
Tetraphlebia C. & R. Felder, 1867
Thiemeia Weymer, 1912

Melanargiina Wheeler, 1903

Melanargia Meigen, 1828
- = Agapetes Billberg, 1820
- = Arge Hübner, 1819 non Arge Schrank, 1802 (Hymenoptera)
- = Ledargia Houlbert, 1922
- = Epimede Houlbert, 1922
- = Parce Oberthür & Houlbert, 1922
- = Halimede Oberthür & Houlbert, 1922
- = Lachesis Oberthür & Houlbert, 1922
- = Argeformia Verity, 1953

Satyrina

Satyrus Latreille, 1810
Arethusana de Lesse, 1951
Argestina Riley, 1923
Aulocera Butler, 1867
Austroypthima Holloway, 1974
Berberia de Lesse, 1951
Boerebia Prout, 1901
- = Erebomorpha Elwes, 1899 non Erebomorpha Walker, 1860 (Geometridae)
Brintesia Fruhstorfer, 1911
- = Oreas Hübner, 1806
- = Oreas Oken, 1815
Callerebia Butler, 1867
Cassionympha van Son, 1955
Chazara Moore, 1893
- = Philareta Moore, 1893 non Philareta Moore, 1881 (Noctuidae)
Coenyra Hewitson, 1865
Coenyropsis van Son, 1958
Davidina Oberthür, 1879
- = Leechia Röber, 1907 non Leechia South, 1901 (Crambidae), nec Leechia Tutt, 1907 (Lycaenidae)
- = Sinosatyrus Lee
Hipparchia Fabricius, 1807
- = Eumenis Hübner, 1819
- = Nytha Billberg, 1820
- = Melania Sodoffsky, 1837
- = Pseudotergumia Agenjo, 1947
- = Neohipparchia de Lesse, 1951
- = Parahipparchia Kurdna, 1977
- = Euhipparchia Kudrna, 1977
Kanetisa Moore, 1893
Karanasa Moore, 1893
Loxerebia Watkins, 1925
- = Hemadara Moore, 1893
Mashuna van Son, 1955
- = Ypthimorpha Overlaet, 1955
Melampias Hübner, 1819
Minois Hübner, 1819
Neita van Son, 1955
Neocoenyra Butler, 1886
Neominois Scudder, 1875
Oeneis Hübner, 1819
- = Chionobas Boisduval, 1833
Paralasa Moore, 1893
Paroeneis Moore, 1893
Physcaeneura Wallengren, 1857
- = Periplysia Gerstaecker, 1871
Pseudochazara de Lesse, 1951
Pseudonympha Wallengren, 1857
Strabena Mabille, 1877
- = Callyphthima Butler, 1880
Stygionympha van Son, 1955
Ypthima Hübner, 1818
- = Xois Hewitson, 1865
- = Kolasa Moore, 1893
- = Thymipa Moore, 1893
- = Nadiria Moore, 1893
- = Pandima Moore, 1893
- = Lohana Moore, 1893
- = Dallacha Moore, 1893
- = Shania Evans, 1912
- = Sundaypthima Uemura, 1996
Ypthimomorpha van Son, 1955

incertae sedis

Dyndirus Capronnier, 1874
Setodocis Billberg, 1820 (genus dubium)

Stamboom

Het onderstaande cladogram geeft de verwantschappen tussen de subtribus van deze geslachtengroep aan zoals die volgens Marin et al. (2011) moeten worden gezien:^[5]

Satyrini

	Parargina

	Lethina

	Mycalesina

	Eritina

	Ragadiina

	Coenonymphina + Hypocystina

	Euptychiina

	Erebiina

	Maniolina

	Yptimina

	Pronophilina

	Melanargiina

	Satyrina

Bronnen, noten en/of referenties

↑ Peña, C., N. Wahlberg, E. Weingartner, U. Kodandaramaiah, S. Nylin, A.V.L. Freitas & A.V.Z. Brower (2006). Higher level phylogeny of Satyrinae butterflies (Lepidoptera: Nymphalidae) based on DNA sequence data. Molecular Phylogenetics and Evolution 40: 29-49.
↑ The higher classification of Nymphalidae, The Nymphalidae Systematics Group.
↑ Een asterisk betekent dat dit geslacht geen deel uitmaakte van het fylogenetisch onderzoek door Pena et al. (2006).
↑ ^a ^b Door Peña (2006) in Coenonymphina geplaatst
↑ Marin, M.A., Peña, C., Freitas, A.V.L., Wahlberg, N. & Uribe, S.I. (2011). From the phylogeny of the Satyrinae butterflies to the systematics of Euptychiina (Lepidoptera, Nymphalidae): history, progress and prospects. Neotropical Entomology 40: 1-13.

[1] Peña, C., N. Wahlberg, E. Weingartner, U. Kodandaramaiah, S. Nylin, A.V.L. Freitas & A.V.Z. Brower (2006). Higher level phylogeny of Satyrinae butterflies (Lepidoptera: Nymphalidae) based on DNA sequence data. Molecular Phylogenetics and Evolution 40: 29-49.

[2] The higher classification of Nymphalidae, The Nymphalidae Systematics Group.

[NotPena-3] Een asterisk betekent dat dit geslacht geen deel uitmaakte van het fylogenetisch onderzoek door Pena et al. (2006).

[coenonymphina-4] Door Peña (2006) in Coenonymphina geplaatst

[Marin2011-5] Marin, M.A., Peña, C., Freitas, A.V.L., Wahlberg, N. & Uribe, S.I. (2011). From the phylogeny of the Satyrinae butterflies to the systematics of Euptychiina (Lepidoptera, Nymphalidae): history, progress and prospects. Neotropical Entomology 40: 1-13.

[1]

[2]

[3]

[4]

[5]